Add the citation: paper link under the title, BibTeX in the app and the README

README.md

@@ -194,3 +194,20 @@ change is resolvable; the demo does this by construction, with the seed fixed at
 
 Measured on the cluster stack (torch 2.4.1); the Space runs torch 2.8.0, which is
 not expected to change these but has not been compared directly.
+
+---
+
+## Citation
+
+If you use CELL-FM, please cite:
+
+```bibtex
+@article{zheng2026virtual,
+  title   = {Virtual experiments bridge sequence and microscopy with generative models},
+  author  = {Zheng, Dihan and Hong, Kibeom and Huang, Bo},
+  journal = {bioRxiv},
+  year    = {2026},
+  doi     = {10.64898/2026.09.13.751243},
+  url     = {https://www.biorxiv.org/content/10.64898/2026.09.13.751243v1}
+}
+```

app.py

@@ -369,12 +369,25 @@ def build_virtual_opencell_section():
                        show_progress="hidden", trigger_mode="always_last")
 
 
+CITATION = """@article{zheng2026virtual,
+  title   = {Virtual experiments bridge sequence and microscopy with generative models},
+  author  = {Zheng, Dihan and Hong, Kibeom and Huang, Bo},
+  journal = {bioRxiv},
+  year    = {2026},
+  doi     = {10.64898/2026.09.13.751243},
+  url     = {https://www.biorxiv.org/content/10.64898/2026.09.13.751243v1}
+}"""
+
+
 with gr.Blocks(title="CELL-FM", theme=gr.themes.Soft(), css=viewer.CSS, head=viewer.HEAD) as demo:
     gr.Markdown(
         """
         # CELL-FM
 
         A virtual microscopy model that bridges microscopy images and protein sequences.
+
+        Paper: Zheng, Hong & Huang, *Virtual experiments bridge sequence and microscopy with
+        generative models*, bioRxiv (2026), [doi:10.64898/2026.09.13.751243](https://www.biorxiv.org/content/10.64898/2026.09.13.751243v1)
         """
     )
 
@@ -384,6 +397,9 @@ with gr.Blocks(title="CELL-FM", theme=gr.themes.Soft(), css=viewer.CSS, head=vie
         with gr.Tab("Virtual OpenCell"):
             build_virtual_opencell_section()
 
+    with gr.Accordion("Citation", open=False):
+        gr.Markdown(f"If you use CELL-FM, please cite:\n\n```bibtex\n{CITATION}\n```")
+
 
 # Download weights and build the models at startup, on CPU. Under ZeroGPU the GPU
 # budget only covers the decorated call, so this must not happen inside it.